九州大学数理生物学研究室

メンバー紹介

巌佐 庸(いわさ よう)

巌佐 庸(いわさ よう)

所属:九州大学理学研究院生物科学部門数理生物学教室、九州大学高等研究院長

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役職:教授

所属学会:日本数理生物学会、日本進化学会、日本生態学会、個体群生態学会、日本動物行動学会、日本分子生物学会、Ecological Society of America、International Society of Behavioral Ecology、Society for Mathematical Biology

連絡先:yohiwasa[at]kyudai.jp
※[at]を@に置き換えて下さい。

数理生物学について

細胞、脳・神経、免疫・感染症、発生、行動、生態、進化などさまざまな生命現象に対して、基本概念を数理モデルとして定式化しその解析やコンピュータシミュレーションによって生命現象を把握するという数理的アプローチが展開されています。これらは数理生物学もしくは理論生物学と呼ばれます。

数理生物学は、ちょうど物理学における理論物理学の役割を、生命科学において果たすものです.生命現象を解明する目的で、数理的手法およびコンピュータシミュレーシ ョンによって行なう研究であれば、なんでも数理生物学の範囲に入るのです。

九州大学の数理生物学研究室について

九州大学理学部の数理生物学研究室は、そのような公式の名前をもつ日本で唯一の研究室として、博士課程修了者は全国の大学の教員や国公立研究機関の研究員として活躍しています。とくに分子進化学、生態学、理論発生学、数理社会学などの分野では、九州大学の数理生物学研究室の出身者の多くが日本での中心的研究者となっています。また米国、ヨーロッパ、オーストラリアをはじめとして様々な国でも活躍しています。

略歴

1975年3月 京都大学理学部卒業
1979年4月~1980年3月 日本学術振興会博士課程奨励研究員(分野:基礎生物学)
1980年4月~1981年9月 日本学術振興会奨励研究員(特定分野:基礎生物学)
1981年10月~1983年10月 米国スタンフォード大学生物科学科博士研究員(Jonathan Roughgarden 教授)
1982年3月 京都大学大学院理学研究科博士課程修了(主査:寺本英教授)
1982年3月 京都大学理学博士取得
1983年11月~1985年1月 米国コーネル大学生態系研究センター研究員(所長:Simon A. Levin 教授)
1985年1月~1992年2月 九州大学理学部助手(講座主任:松田博嗣教授)
1992年3月~1999年3月 九州大学理学部教授(生物学教室数理生物学講座担当)
1999年4月~2000年3月 九州大学大学院理学研究科教授
2000年4月~現在 九州大学理学研究院教授(生物科学部門動態生物学講座担当、数理生物学研究室)
2010年4月~現在 九州大学高等研究院長

業績

(269) Liao, Jinbao; Li, Zhenqing; Hiebeler, David; Iwasa, Yoh; Bogaert, Jan; Nijs, Ivan. 2013. Species persistence in landscapes with spatial variation in habitat quality: a pair approximation model. Journal of Theoretical Biology (in press)
(268) Yamaguchi, S., K. Sawada, Y. Yusa, and Y. Iwasa. 2013. Dwarf males and hermaphrodites can coexist in marine sedentary species if the opportunity to become a dwarf male is limited. Journal of Theoretical Biology (in press)
(267) Tachiki, Y., and Y. Iwasa. 2013. Coevolution of mast seeding in trees and extended diapause of seed predators. Journal of Theoretical Biology (in press)
(266) Iwasa, Y. and J-H. Lee. 2013. Graduated punishment is efficient in resource management if people are heterogeneous. Journal of Theoretical Biology (in press)
(265) Haeno, H., Y.E. Maruvka, Y. Iwasa, and F. Michor. 2013. Stochastic tunneling of two mutations in a population of a constant number of cells. PLoS ONE (in press)
(264) Yamaguchi, R. and Y. Iwasa. 2013. Reproductive character displacement by the evolution of female mate choice. Evolutionary Ecology Research 15(1): 25-41.
(263) Fukano, Y., Y. Tachiki, T. Yahara, and Y. Iwasa. 2013. Soil disturbances may suppress the invasion of alien plants under plant-soil-feedback. Ecological Modelling 260:42-49.
(262) Yahara, T. F. Javadi, Y. Onoda, L. P. de Queiroz, D. P. Faith, D. E. Prado, M. Akasaka, T. Kadoya, F. Ishihama, S. Davies, J. W. F. Slik, T.Yi, K. Ma, B. Chen, D. Darnaedi, R. T. Pennington, M. Tuda, M. Shimada, M. Ito, A. N. Egan, S. Buerki, N. Raes, T. Kajita, M. Mimura, H. Tachida, Y. Iwasa, G. F. Smith, J. E. Victor, & T. Nkonki. 2013. Global legume diversity assessment: concepts, key indicators, and strategies. Taxon 62:249-266.
(261) Yamaguchi, S., K. Sawada, Y. Yusa, and Y. Iwasa. 2013. Dwarf males, hermaphrodites, and large females in marine species: a dynamic optimization model of sex allocation and growth. Theoretical Population Biology 85: 49-57.
(260) Halley, J.M., D., Vokou, and Y. Iwasa. 2013. Comennto on "Extinction debt and windows of conservation opportunity in the Brazilian Amazon". Science 339:271.
(259) Seto, M., N. Takamura, and Y. Iwasa, 2013. Individual and combined suppressive effects of submerged and floating-leaved macrophytes on algal blooms. Journal of Theoretical Bilogy 319: 122-133.
(258) Wakano, J., and Y. Iwasa. 2013. Evolutionary branching in a finite population: deterministic branching versus stochastic branching. Genetics 193: 229-241.
(257) Koizumi, Y., Y. Iwasa, and T. Hirashima. 2012. Mathematical study of the role of Delta/Notch inhibition during primary branching of Drosophila Trachea development. Biophisical Journal 103:2549-1559.
(256) Halley, J.M., and Y. Iwasa. 2012. Neutrality without incoherence: a response to Clark. Trends in Ecology and Evolution 27: 363.
(255) Lee, J.H. and Y. Iwasa. 2012. Optimal investment in enhancing social concern on biodiversity conservation: a dynamic approach. Theoretical Population Biology 82:177-186.
(254) Tanaka, C.M. and Y. Iwasa. 2012. Cultural evolution of a belief controlling human mate choice: dynamic modeling of the hinoeuma superstition in Japan. Journal of Theoretical Bilogy. 309:20-28.
(253) Takashina, N. A. Mougi, and Y. Iwasa, Paradox of Marine Protected Area: Suppression of Fishery May Cause Species Loss. Population Ecology 54: 475-485.
(252) Bessho, K. and Y. Iwasa. 2012. Variability in the evolutionarily stable seasonal timing of germination and maturation and the mode of competition. Journal of Theoretical Biology 304:66-80.
(251) Satake, A. and Y. Iwasa. 2012. A stochastic model of chromatin modification: cell population coding of winter memory in plants. Journal of Theoretical Biology 302:6-17.
(250) Tachiki, Y. and Y. Iwasa, 2012. Evolutionary jumping and breakthrough in the trees' masting evolution. Theoretical Population Biology 81:20-31.
(249) Hironaka, K., Y. Iwasa, and Y. Morishita. 2012. Multiple feedback loops achieve robust localization of wingless expression in Drosophila notum development. Journal of Theoretical Biology 292:18-29.
(248) Uchinomiya, K and Y. Iwasa. 2011. Burrowing in the bark or in the sapwood: a dynamic game between a mother and her offspring. Evolutionary Ecology Researech 13:607-623.
(247) Morishita, Y. and Y. Iwasa. 2011. Coding design of positional information for robust morphogenesis. Biophysical Journal 101:2324-2335.
(246) Lee, Joung-Hun and Y. Iwasa. 2011. Tourists and traditional divers in a common fishing ground. Ecological Economics 70:2350-2360.
(245) Muko, S., and Y. Iwasa. 2011. Long-term effect of coral transplantation: Restoration-goals and the choice of species. Journal of Theoretical Biology 280:127-138.
(244) Mougi, A., and Y. Iwasa. 2011. Green world maintained by adaptation. Theoretical Ecology 4: 201-210.
(243) Mougi, A., O. Kishida, and Y. Iwasa. 2011. Coevolution of phenotypic plasticity in predator and prey: why inducible offense is rarer than inducible defense? Evolution 65: 1079-1081.
(242) Iwasa, Y., and F. Michor. 2011. Evolutionary dynamics of tumor diversity. PLoS ONE 6: e17866.
(241) Mougi, A. and Y. Iwasa. 2011. Unique coevolutionary dynamics in a predator-prey system. Journal of Theoretical Biology 277:83-89.
(240) Saeki, K. and Y. Iwasa. 2011. T cell anergy as a strategy to reduce the risk of autoimmunity. Journal of Theoretical Biology 277:74-82.
(239) Halley, J.M., and Y. Iwasa. 2011. Neutral theory as a predictor of avifaunal extinctions following habitat loss. Proc. Nat. Acad. Sci. USA 108:2316-2321.
(238) Muko, S., and Y. Iwasa. 2011. Optimal choice of species and size classes for transplantation of coral community. Journal of Theoretical Biology 273:130-137.
(237) Seto, M., and Y. Iwasa. 2011. Regime shift and robustness of organism-created environment: a model for microbial ecosystem. Journal of Theoretical Biology 269:297-306.
(236) Bessho, K. and Y. Iwasa. 2010. Optimal seasonal schedule of heteromorphic and isomorphic life-cycles in macroalgae, and their relative dominance. Journal of Theoretical Biology 267:201-212.
(235) Tachiki, Y., Y. Iwasa, and A. Satake. 2010. Pollinator coupling can induce synchronized flowering among different plant species. Journal of Theoretical Biology 267:153-163.
(234) Tachiki, Y. and Y. Iwasa. 2010. Both seedling banks and specific seed predators promote the evolution of synchronized and intermittent reproduction (masting) in trees. Journal of Ecology 98:1398-1408.
(233) Brooks, M., and Y. Iwasa. 2010. Size-dependent sex change is the ESS without size-advantage of reproduction. Theoretical Population Biology 78:183-191.
(232) Mougi, A. and Y. Iwasa. 2010. Evolution towards oscillation or stability in a predator-prey system. Proceeding of Royal Society London ser. B 277: 3163-3171.
(231) Uriu, K., Y. Morishita, and Y. Iwasa. 2010. Synchronized oscillation of the segmentation clock gene in vertebrate development. Journal of Mathematical Biology 61:217-229.
(230) Uriu, K., Y. Morishita, and Y. Iwasa. 2010. Synchronization of segmentation clock is promoted by random cell movement. Proc. Nat. Acad. Sci. USA 107:4979-4984.
(229) Tazzyman, S.J., and Y. Iwasa. 2010. Sexual selection can increase the effect of random genetic drift -- A quantitative genetic model of polymorphism in O. Pumilio, the Strawberry poison-dart frog. Evolution 64:1719-1728.
(228) Iwasa, Y., Y. Suzuki-Ohno, and H. Yokomizo. 2010. Paradox of nutrient removal in coupled socio-economic and ecological dynamics for lake water pollution Theoretical Ecology 3:113-122.
(227) Saeki. K. and Y. Iwasa. 2010. Optimal number of regulatory T cells. Journal of Theoretical Biology 263:210-218.
(226) Uehara, T. and Y. Iwasa. 2010. Global mutations and local mutations have very different effects on evolution, illustrated by mixed strategies of asymmetric binary games. Journal of Theoretical Biology 262: 223-231.
(225) Hirashima, T., Y. Iwasa, and Y. Morishita. 2009. Mechanisms for split localization of Fgf10 observed in early lung development. Developmental Dynamics 238:2813–2822.
(224) Saeki, K., and Y. Iwasa. 2009. Advantage of having regulatory T cells requires localized suppression of immune reactions. Journal of Theoretical Biology 260: 392-401.
(223) Bessho, K., and Y. Iwasa. 2009. Heteromorphic and isomorphic alternations of generations in macroalgae, as adaptations to a seasonal environment. Evolutionary Ecology Research 111: 691-771.
(222) Morishita, Y. and Y. Iwasa, 2009. Accuracy of positional information provided by multiple chemical gradients with correlated noises. Physical Review E 79: art. no. 061905.
(221) Gokhale, C., Y. Iwasa, M.A. Nowak, and A. Traulsen. 2009. The pace of evolution across fitness valleys. Journal of Theoretical Biology 259: 613-620.
(220) Hirashima, T., Y. Iwasa, and Y. Morishita. 2009. Dynamic modeling of ureteric tube branching in the early kidney development. Journal of Theoretical Biology 259: 58-66.
(219) Suzuki, Y., and Y. Iwasa. 2009. The coupled dynamics of human socio-economic choice and lake water system: the interaction of two sources of nonlinearity. Ecological Research 24: 479-489.
(218) Uriu, K., Y. Morishita and Yoh Iwasa. 2009. Traveling wave formation in vertebrate segmentation. Journal of Theoretical Biology 257: 385-396.
(217) Suzuki, Y., and Y. Iwasa, 2009. Conflict between groups of players in coupled socio-economic and ecological dynamics. Ecological Economics 68: 1006-1115
(216) Morishita, Y. and Y. Iwasa, 2009. Estimating the spatio-temporal pattern of volumetric growth rate from fate maps in chick limb development. Developmental Dynamics 238: 415-422.
(215) Ohtsuki, H., Y.Iwasa, and M.A. Nowak. 2009. Indirect reciprocity provides only a narrow margin of efficiency for the costly punishment. Nature 457:179-182.
(214) Satake, A., S.A. Levin and Y. Iwasa. 2008. Comparison between perfect information and passive-adaptive social learning models of forest harvesting. Theoretical Ecology 1:189-197.
(213) Tachiki, Y. and Y. Iwasa. 2008. Role of gap dynamics in the evolution of masting of trees. Evolutionary Ecology Research 10: 893-905.
(212) Morishita, Y. and Y. Iwasa. 2008. Growth based morphogenesis in vertebrate limb bud. Bulletin of Mathematical Biology 70, 1957-1978.
(211) Morishita, Y. and Y. Iwasa. 2008. Optimal placement of multiple morphogen sources. Physical Review E 77: 41909
(210) Muko, S. and Y. Iwasa. 2008. Spatial heterogeneity of mortality and temporal fluctuation in fertility promote coexistence but not vice versa: a random-community approach. Journal of Theoretical Biology 253:503-600.
(209) Hirashima, T., Y. Iwasa, and Y. Morishita. 2008. Distance between AER and ZPA is defined by feed-forward loop and is stabilized by their feedback loop in vertebrate limb bud. Bulletin of Mathematical Biology 70:438-459.
(208) Haeno, H., Y. Iwasa, and F. Michor. 2007. The evolution of two mutations during clonal expansion. Genetics 177: 2209-2221.
(207) Uriu, K. and Y. Iwasa. 2007. Turing pattern formation with two kinds of cells and a diffusive chemical. Bulletin of Mathematical Biology 69: 2515-2536
(206) Kon, R. and Y. Iwasa, 2007. Single class orbits in nonlinear Leslie matrix models for semelparous populations. Journal of Mathematical Biology 55: 781-802.
(205) Traulsen, A., Y. Iwasa, and M.A. Nowak. 2007. The fastest evolutionary trajectory. J. Theoretical Biology 249: 617-623.
(204) Taylor, C., J. Chen, and Y. Iwasa. 2007. Cooperation maintained by fitness adjustment. Evolutionary Ecology Research 9: 1023-1041.
(203) Satake, A., M. Janssen, S.A. Levin, and Y. Iwasa. 2007. Synchronized deforestation induced by social learning under uncertainty of forest-use value Ecological Economics 63:452-462.
(202) Uehara, T., Y. Iwasa, and H. Ohtsuki. ESS distribution of display duration in animal contests to assess an opponent before fighting or fleeing. Evolutionary Ecology Research 9:395-408.
(201) Satake, A., H.M. Leslie, Y. Iwasa, and S.A. Levin. 2007. Coupled ecological-social dynamics in a forested landscape: spatial interactions and information flow. Journal of Theoretical Biology 246:695-707.
(200) Shudo, E., J. Yang, A. Yoshimura, and Y. Iwasa. 2007. Robustness of the signal transduction system of the mammalian JAK/STAT pathway and dimerization steps. Journal of Theoretical Biology 246:1-9.
(199) Iwasa, Y., T. Uchida, and H. Yokomizo. 2007. Nonlinear behavior of the socio-economic dynamics for lake water pollution control. Ecological Economics 63:219-229.
(198) Schlicht, R., and Y. Iwasa. 2007. Spatial pattern analysis in forest dynamics: Deviation from power law and direction of regeneration waves. Ecological Research 22: 197-203.
(197) Haeno, H. and Y. Iwasa. 2007. Probability of resistance evolution in the host by the exponentially growing virus. Journal of Theoretical Biology 246: 323-331.
(196) Ohtsuki, H. and Y. Iwasa. 2007. Global analysis of evolutionary dynamics and exhaustive search for social norms that maintain cooperation by reputation. Journal of Theoretical Biology 244:518-531.
(195) Yokomizo, H., P. Haccou and Y. Iwasa. 2007. Optimal conservation strategy in fluctuating environments with species interactions: resources enhancement of the native species versus extermination of the alien species. Journal of Theoretical Biology 244:46-58.
(194) Iwasa, Y. and G. Wada. 2006. Complex life cycle and the body sizes at life history transitions for macroparasites. Evolutionary Ecology Research 8:1427-1443.
(193) Taylor, C., Y. Iwasa, and M.A. Nowak. 2006. A symmetry of fixation time in evolutionary dynamics. Journal of Theoretical Biology 243:245-251.
(192) Feugier, G.F. and Y. Iwasa. 2006, How canalization can make loops: a new model of reticulated leaf vascular pattern formation. Journal of Theoretical Biology 243:235-244.
(191) Michor, F., Y. Iwasa and M.A. Nowak. 2006. The age incidence of chronic myeloid leukemia can be explained by a one-mutation model. Proc. Natl. Acad. Sci. USA 103:14931-14934.
(190) Schlict, R. and Y. Iwasa. 2006. Deviation from power-law, spatial data of forest canopy gaps, and three lattice models Ecological Modelling 198:399-408.
(189) Kurosawa, G., K. Aihara, and Y. Iwasa. 2006. A model for circadian rhythm of cyanobacteria, which maintains oscillation without gene expression. Biophysical Journal 91: 2015-2025.
(188) Schlicht, R. and Y. Iwasa. 2006. Direction of regeneration waves in grid-based models for forest dynamics. Journal of Theoretical Biology 242:365-371.
(187) Nakamaru, M. and Y. Iwasa. 2006. The coevolution of altruism and punishment: the role of selfish punishers. Journal of Theoretical Biology 240:475-488.
(186) Michor, F. and Y. Iwasa. 2006. Dynamics of metastasis suppressor gene inactivation. Journal of Theoretical Biology 241:676-689.
(185) Nowak, M.A., F. Michor, and Y. Iwasa. 2006. Genetic instability and clonal expansion. Journal of Theoretical Biology 241: 26-32.
(184) Michor, F., M.A. Nowak, and Y. Iwasa. 2006. Stochastic dynamics of metastasis formation. Journal of Theoretical Biology 240: 521-530.
(183) Satake, S. and Y. Iwasa. 2006. Coupled ecological and social dynamics in a forested landscape: the deviation of the individual decisions from the social optimum. Ecological Research 21: 370-379.
(182) Iwasa, Y., M.A. Nowak, and F. Michor. 2006. Evolution of resistance in clonal expansion. Genetics 172:2557-2566.
(181) Zea-Cabrera, E., Y. Iwasa, S. Levin, I. Rodriguez-Iturbe, 2006. Tragedy of the commons of plant water use. Water Resources Research 42: W06D02, doi:10.1029/2005WR004514.
(180) Ohtsuki, H. and Y. Iwasa. 2006. The leading eight: social norms that can maintain cooperation by indirect reciprocity. Journal of Theoretical Biology 239:435-444.
(179) Mochizuki, A., K. Yahara, Kobayashi, I., and Y. Iwasa. 2006. Selfish gene's strategy for symbiosis in the genome. Genetics 172: 1309-1323.
(178) Michor, F., M.A. Nowak and Y. Iwasa. 2006. Evolution of resistance to cancer therapy. Current Pharmaceutical Design 12:261-271.
(177) Nakamaru, M. and Y. Iwasa. 2005. The evolution of altruism by costly punishment in the lattice structured population: score-dependent viability versus score-dependent fertility. Evolutionary Ecology Research 7: 853-870.
(176) Michor, F., Y. Iwasa, B. Vogelstein, C. Lengauer, and M.A. Nowak. 2005. Dynamics of colorectal cancer. Seminars in Cancer Biology 15: 484-493.
(175) Shoji, H. and Y. Iwasa. 2005 Labyrinth versus straight-stripes generated by two-dimensional Turing systems. Journal of Theoretical Biology 237: 104-116.
(174) Joron, M. and Y. Iwasa. 2005. The evolution of a Muellerian mimic in a spatially-distributed community. Journal of Theoretical Biology 237: 87-103.
(173) Feugier, F.G., A. Mochizuki and Y. Iwasa, 2005. Self-organizing formation of vascular system of plant leaves: co-orientation between auxin flux and pump proteins. Journal of Theoretical Biology 236: 366-375.
(172) Michor, F., T.P. Hughes, Y. Iwasa, S. Branford, N.P. Shah, C.L. Sawyers, and M.A. Nowak. 2005. Dynamics of chronic myeloid leukemia. Nature 435:1267-1270.
(171) Uehara, T., H. Yokomizo, and Y. Iwasa. 2005. Mate-choice copying as Bayesian decision making. American Naturalist 165:403-410.
(170) Kurosawa, G., and Y. Iwasa, 2005. Temperature compensation in circadian clock models. Journal of Theoretical Biology 233: 453-468.
(169) Irie, T., and Y. Iwasa. 2005. Optimal growth pattern of defensive organs: the diversity of shell growth among Molluscs. American Naturalist 165, 238-249.
(168) Iwasa, Y., F. Michor, N.L. Komarova, and M.A. Nowak. 2005. Population genetics of tumor suppressor genes. Journal of Theoretical Biology 233: 15-23.
(167) Michor, F., Y. Iwasa, B. Vogelstein, C. Lengauer, and M.A. Nowak. 2005. Can chromosomal instability initiate tumorigenesis? Seminars in Cancer Biology 15: 43-39.
(166) Hakoyama, H. and Y. Iwasa. 2005. Extinction risk of a meta-population: aggregation approach. Journal of Theoretical Biology 232: 203-216.
(165) Iwasa, Y., F. Michor, and M. A. Nowak.2005. Virus evolution within patients increases pathogenicity. Journal of Theoretical Biology 232:17-26.
(164) Wodarz, D., N. Komarova, and Y. Iwasa. 2004. On the emergence of multifocal cancers. Journal of Carcinogenesis 3: 13. doi:10.1186/1477-3163-3-13.
(163) Ohtsuki, H. and Y. Iwasa. 2004. How should we define goodness? -- reputation dynamics in indirect reciprocity. Journal of Theoretical Biology 231:107-120.
(162) Gratzer, G., C. Canham, U. Dieckmann, A. Fischer, Y. Iwasa, R., Law, M. Lexer, H. Sandman, T. Spies, B. Splechtna, and J. Szwagrzyk. 2004. Spatio-temporal development of forests--current trends in field studies and model. Oikos 107: 3-15.
(161) Yokomizo, H., P. Haccou and Y. Iwasa. 2004. Multiple-year optimization of conservation effort and monitoring effort for a fluctuating population. Journal of Theoretical Biology 230:157-171.
(160) Schlicht, R. and Y. Iwasa. 2004. Forest gap dynamics and the Ising model. Journal of Theoretical Biology 230: 65-75.
(159) Nowak, M.A., F. Michor, N.L. Komarova and Y. Iwasa. 2004. Evolutionary dynamics of tumor suppressor gene inactivation. Proc. Natl. Acad. Sci. USA 101: 10635-10638.
(158) Iwasa, Y., F. Michor, and M.A. Nowak. 2004. Some basic properties of immune selection. Journal of Theoretical Biology 229: 179-188.
(157) Iwasa, Y., F. Michor and M.A. Nowak. 2004. Stochastic tunnels in evolutionary dynamics. Genetics 166:1571-1579.
(156) Shudo, E. and Y. Iwasa. 2004 Dynamic optimization of defense, immune memory, and post-infection pathogen levels in mammals. Journal of Theoretical Biology 228:17-29.
(155) Michor, F., Y. Iwasa, H. Rajagopalan, C. Lengauer, M. A. Nowak. 2004. A linear model of colon cancer initiation. Cell Cycle 3:358-362.
(154) Iwasa, Y. and A. Satake. 2004. Mechanisms inducing spatially extended synchrony in mast seeding: the role of pollen coupling and environmental fluctuation Ecological Research 19: 13-20.
(153) Michor, F., Y. Iwasa, and M.A. Nowak. 2004. Dynamics of cancer progression. Nature Reviews Cancer 4:197-205.
(152) Satake, A., Y. Iwasa, H. Hakoyama, and S.P. Hubbell. 2004. Estimating local interaction from spatio-temporal forest data, and Monte Carlo Bias Correction. Journal of Theoretical Biololgy 226:225-235.
(151) Iwasa, Y., F. Michor, and M. Nowak. 2004. Evolutionary dynamics of invasion and escape. Journal of Theoretical Biology 226:205-214.
(150) Hakoyama, H. and Y. Iwasa. 2004. Coexistence of a sexual and a unisexual forms by parasites. Journal of Theoretical Biology 226:185-194.
(149) Irie, T. and Y. Iwasa. 2004. Optimal growth model for the latitudinal cline of shell morphology in cowries (genus Cypraea). Evolutionary Ecology Research 5:1133-1149.
(148) Iwasa, Y., F. Michor, and M. Nowak. 2003. Evolutionary dynamics of escape from biomedical intervention. Proceeding of Royal Society London ser B. 270:2573-2578.
(147) Nowak, M.A., F. Michor, and Y. Iwasa. 2003. The linear process of somatic evolution. Proc. Nat. Acad. Sci. USA 100:14966-14969.
(146) Michor, F., S. A. Frank, R. M. May, and Y. Iwasa, and Nowak, M.A. 2003. Somatic selection for and against cancer. Journal of Theoretical Biology 225:377-382.
(145) Michor, F., M.A. Nowak, S.A. Frank, and Y. Iwasa. 2003. Stochastic elimination of cancer cells. Proceeding of Royal Society London ser. B. 270: 2017-2024.
(144) Muko, S. and Y. Iwasa. 2003. Incomplete mixing promotes species coexistence in a lottery model with permanent spatial heterogeneity. Theoretical Population Biology 64:359-368.
(143) Shoji, H., Y. Iwasa, and S. Kondo. 2003. Stripes, Spots or Reversed Spots in Two-dimensional Turing Systems. Journal of Theoretical Biology 224:339-350.
(142) Nakanishi, J., M. Gamo, Y. Iwasa, and Y. Tanaka. 2003. Environmental Risk Evaluation of Chemicals: achievements of the project and seeds for future --- Development of metrics for evaluating risks. Chemosphere 53:389-398.
(141) Nakamaru, M., Y. Iwasa, and J. Nakanishi. 2003. Extinction risk to bird populations caused by DDT exposure. Chemosphere 53:377-387.
(140) Yokomizo, H., P. Haccou and Y. Iwasa. 2003. Conservation effort for a population in a fluctuating environment and assessment of the population size. Journal of Theoretical Biology 224:167-182.
(139) Yokomizo, H., J. Yamashita, and Y. Iwasa. 2003. Optimal conservation effort for a population in a stochastic environment. Journal of Theoretical Biology 220:215-231.
(138) Shoji, H. and Y. Iwasa. 2003. Pattern selection and the direction of stripes in two-dimensional Turing systems for skin pattern formation of fishes. Forma 18:3-18.
(137) Shudo, E., P. Haccou, and Y. Iwasa 2003. Optimal choice between feedforward and feedback control in gene expression to cope with unpredictable danger. Journal of Theoretical Biology 223:149-160.
(136) Shoji, H., A. Mochizuki, Y. Iwasa, M. Hirata, T. Watanabe, S. Hioki, and S. Kondo. 2003. Origin of directionality in the fish stripe pattern. Developmental Dynamics 226:627-633.
(135) Frank, S., Y. Iwasa, and M. Nowak. 2003. Pattern of cell division and the risk of cancer. Genetics 163:1527-1532.
(134) Takano, R., A. Mochizuki, and Y. Iwasa. 2003. Possibility of tissue separation caused by cell adhesion. Journal of Theoretical Biology 221:459-474.
(133) Michor, F., Y. Iwasa, N. L. Komarova, and M. A. Nowak. 2003. Local regulation of homeostasis favors chromosomal instability. Current Biology 13:581-584.
(132) Tohya, S., A. Mochizuki, and Y. Iwasa. 2003. Random cell rearrangement can form cone mosaic patterns in fish retina and explain the difference between zebrafish and medaka. Journal of Theoretical Biology 221:289-300.
(131) Iwasa, Y. and A. Satake. 2002. Synchronized reproduction of trees caused by pollen exchange: modeled as coupled chaos systems. Acta Biologica Venezuelica 22: 45-57.
(130) Kurosawa, G., and Y. Iwasa. 2002. Saturation of enzyme kinetics in circadian clock models. Journal of Biological Rhythms 17:568-588.
(129) Satake, A. and Y. Iwasa. 2002 The synchronized and intermittent reproduction of forest trees is mediated by the Moran effect, only in association with pollen coupling. Journal of Ecology 90:830-833.
(128) Shudo, E. and Y. Iwasa. 2002. Optimal defense strategy: storage versus new production. Journal of Theoretical Biology 219:309-323.
(127) Kurosawa, G., A. Mochizuki, and Y. Iwasa. 2002. Processes promoting oscillations -- comparative study of circadian clock models. Journal of Theoretical Biology 216:193-208.
(126) Satake, A. and Y. Iwasa. 2002. Spatially limited pollen exchange and a long range synchronization of forest trees. Ecology 83:993-1005.
(125) Shoji, H.,Y. Iwasa, A. Mochizuki, and S. Kondo. 2002. Directionality of stripes formed by anisotropic reaction-diffusion models. Journal of Theoretical Biology 214:549-561.
(124) Nakamaru, M., Y. Iwasa and J. Nakanishi. 2002. Extinction risk of DDT to herring gull (Larus argentatus) populations. Environmental Toxicology and Chemistry 21:195-202.
(123) Satake, A., A. Sasaki, and Y. Iwasa. 2001. Variable timing of reproduction in unpredictable environments: adaptation of flood plain plants. Theoretical Population Biology 60:1-15.
(122) Iwasa, Y. and A. Pomiankowski. 2001. The evolution of X-linked genomic imprinting. Genetics 158:1801-1809.
(121) Muko, S., K. Sakai and Y. Iwasa. 2001. Size distribution dynamics for a marine sessile organism with space-limitation in growth and recruitment: an application to a coral population. Journal of Animal Ecology 70:579-589.
(120) Muko, S., K. Sakai and Y. Iwasa. 2001. Dynamics of marine sessile organisms with space-limited growth and recruitment: an application to corals. Journal of Theoretical Biology 210:67-80.
(119) Shudo, E. and Y. Iwasa. 2001. Inducible defense against pathogens and parasites: optimal choice among multiple options. Journal of Theoretical Biology 209:233-247.
(118) Post, E., S.A. Levin, Y. Iwasa, and N. C. Stenseth. 2001. Reproductive asynchrony increases with environmental disturbance. Evolution 55:830-834.
(117) Falush, D., E.W. Almqvist, R.R. Brinkmann,Y.Iwasa, and M.R. Hayden. 2001. Measurement of mutational flow implies a high new mutation rate for Huntington disease and substantial under-ascertainment of late onset cases. American Journal of Human Genetics 68: 373-385.
(116) Muko, S. and Y. Iwasa, 2000. Species coexistence by permanent spatial heterogeneity in a lottery model. Theoretical Population Biology 57:273-284.
(115) Iwasa, Y., H. Hakoyama, M. Nakamaru, and J. Nakanishi, 2000. Estimate of population extinction risk and its application to ecological risk management. Population Ecology 42: 73-80.
(114) Hakoyama, H. and Y. Iwasa. 2000. Bias correction and confidence intervals for parametric models based on the Monte-Carlo sampling method. Japanese Journal of Biometrics 20: 143-154.
(113) Hakoyama, H. and Y Iwasa, 2000. Extinction risk of a density-dependent population estimated from a time series of population size. Journal of Theoretical Biology 204:337-359.
(112) Hakoyama, H., Y. Iwasa, and J. Nakanishi. 2000. Comparing risk factors for population extinction. Journal of Theoretical Biology 204:327-336.
(111) Iwasa, Y. 2000. Dynamic optimization of plant growth. Evolutionary Ecology Research 2: 437-455.
(110) Satake, A. and Y. Iwasa, 2000. Pollen-Coupling of forest trees, forming synchronized and periodic reproduction out of chaos. Journal of Theoretical Biology 203:63-84.
(109) Nakamaru, M. and Y. Iwasa. 2000.Competition by allelopathy proceeds in traveling waves: colicin-immune strain aids colicin-sensitive strain. Theoretical Population Biology 57:131-144.
(108) Kamo, M. and Y. Iwasa. 2000. Preference for consonances as a byproduct of training. Evolutionary Ecology Research 2:375-383.
(107) Suzuki, H. and Y. Iwasa. 1999. Estimation of the GA performance in a Babel-like fitness landscape: acceleration effect of crossover. Evolutionary Computation 7:275-310.
(106) Tohya, S., A. Mochizuki and Y. Iwasa. 1999. Formation of cone mosaic of zebrafish retina. Journal of Theoretical Biology 200:231-244.
(105) Iwasa, Y. and A. Pomiankowski. 1999. Good parent and good genes models of handicap evolution. Journal of Theoretical Biology 200:97-109.
(104) Falush, D.P. and Y. Iwasa. 1999. Size dependent mutation and microsatellite constraint. Molecular Biology and Evolution 16:960-966.
(103) Iwasa, Y. and A. Pomiankowski. 1999. Sex specific X chromosome expression caused by genomic imprinting. Journal of Theoretical Biology 197:487-495.
(102) Iwasa, Y., A. Mochizuki and Y. Takeda. 1999. The evolution of genomic imprinting: abortion and overshoot explain aberrations. Evolutionary Ecology Research 1:129-150.
(101) Tuda, M. and Y. Iwasa. 1998. Evolution of contest competition and its effect on host-parasitoid dynamics. Evolutionary Ecology 12:855-870.
(100) Iwasa, Y., M. Nakamaru and S.A. Levin. 1998. Allelopathy of bacteria in a lattice population: competition between colicin-sensitive and colicin-producing strains. Evolutionary Ecology 12:785-802.
(99) Satake, A., T. Kubo, and Y. Iwasa. 1998. Noise induced regularity of spatial wave patterns in subalpine Abies forests. Journal of Theoretical Biology 195:465-479.
(98) Takesue, A., A. Mochizuki and Y. Iwasa.1998. Cell-differentiation rules that generate regular mosaic patterns: modelling motivated by cone mosaic formation in fish retina. Journal of Theoretical Biology 194:575-586.
(97) Nakamaru, M., H. Nogami and Y. Iwasa. 1998. Score-dependent fertility model for the evolution of cooperation in a lattice. Journal of Theoretical Biology 194:101-124.
(96) Tohya, S., A. Mochizuki, S. Imayama and Y. Iwasa, 1998. On rugged shape of skin tumor (Basal cell carcinoma). Journal of Theoretical Biology 194:65-78.
(95) Pomiankowski, A. and Y. Iwasa. 1998. Runaway ornament diversity caused by Fisherian sexual selection. Proc. Nat. Acad. Sci. USA 95(9):5106-5111.
(94) Mochizuki, A., N. Wada, H. Ide and Y. Iwasa. 1998. The cell-cell adhesion in the limb-formation, estimated from photographs of cell sorting experiments based on a spatial stochastic model. Developmental Dynamics 211:204-214.
(93) Iwasa, Y. and Y. Harada. 1998. Female mate preference to maximize paternal care. II. Female competition leads to monogamy. American Naturalist 151:367-382.
(92) Iwasa, Y. 1998. The conflict theory of genomic imprinting: how much can be explained? Current Topics in Developmental Biology 40:255-293.
(91) Halley, J. and Y. Iwasa. 1998. Extinction rate of a population with both demographic and environmental stochasticity. Theoretical Population Biology 53:1-15.
(90) Kamo, M., T. Kubo and Y. Iwasa. 1998. Neural network for female mate preference, trained by a genetic algorithm. Philosophical Transaction Royal Society of London series B 353:399-406.
(89) Haccou, P. and Y. Iwasa. 1998. Robustness of optimal mixed strategies. Journal of Mathematical Biology 36:485-496.
(88) Yamauchi, A., Y. Matsumiya and Y. Iwasa. 1997. The opimal age-dependent sustainable fishery. Researches on Population Ecology 38:139-148.
(87) Ezoe, H. and Y. Iwasa. 1997. Evolution of condition-dependent dispersal: a genetic algorithm search for the ESS reaction norm. Researches on Population Ecology 38:127-137.
(86) Harada, Y., S. Kawano and Y. Iwasa. 1997. Probability of clonal identity: inferring the relative success of sexual vs. clonal reproduction from spatial genetic patterns. Journal of Ecology 85:591-600.
(85) Takenaka, Y., H. Matsuda and Y. Iwasa. 1997. Competition and Evolutionary Stability of Plants in a Spatially Structured Habitats. Researches on Population Ecology 39:67-75.
(84) Klinkhamer, P.G.L., Y. Iwasa, T. Kubo and T. de Jong. 1997. Herbivores and the evolution of the semelparous perennial life-history of plants. Journal of evolutionary Biology 10: 529-550.
(83) Mochizuki, A., Y. Takeda, H. Ide and Y. Iwasa. 1997. A stochastic model for cell sorting and its application. Forma 12:107-122.
(82) Takeda, Y. and Y. Iwasa. 1997. A neural network modeling for concentration-dependent pair-rule gene expression. Forma 12:123-142.
(81) Iwasa, Y. and T. Kubo. 1997. Optimal size of storage for recovery after unpredictable disturbances. Evolutionary Ecology 11:41-65.
(80) Nakamaru, M., H. Matsuda and Y. Iwasa. 1997. The evolution of cooperation in a lattice-structured population. Journal of Theoretical Biology 184:65-81.
(79) Haccou, P. and Y. Iwasa. 1996. Establishment probability in fluctuating environments: a branching process model. Theoretical Population Biology 50:254-280.
(78) Harada, Y. and Y. Iwasa. 1996. Analyses of spatial patterns and population processes of clonal plants. Researches on Population Ecology 38:153-164.
(77) Mochizuki, A., Y. Takeda and Y. Iwasa. 1996. Evolution of genomic imprinting: why are so few genes imprinted ? Genetics 144:1283-1295.
(76) Iwasa, Y., T. Kubo, N. van Dam and T.J. de Jong, 1996. Optimal level of chemical defense decreasing with leaf age. Theoretical Population Biology 50:124-148.
(75) Kubo, T., Y. Iwasa and N. Furumoto, 1996. Forest spatial dynamics with gap expansion: total gap area and gap size distribution. Journal of Theoretical Biology 180:229-246.
(74) Mochizuki, A., Y. Iwasa and Y. Takeda, 1996. A stochastic model for cell sorting and measuring cell-cell adhesion. Journal of Theoretical Biology 179:129-146.
(73) Harada, Y. and Y. Iwasa. 1996. Female mate preference to receive maximum paternal care: a two-step game. American Naturalist 147:996-1027.
(72) Kunin, W. and Y. Iwasa, 1996. Pollinator foraging strategies in mixed floral arrays: density effects and floral constancy. Theoretical Population Biology 49:232-263.
(71) van Dam, N., T.J. de Jong, Y. Iwasa and T. Kubo, 1996. Optimal distribution of pyrrolizidine alkaloids in rosette plants of Cynoglossum officinale L.: are plants smart investors ? Functional Ecology 10:128-136.
(70) Andersson, M., and Y. Iwasa, 1996. "Sexual selection". Trends in Ecology and Evolution 11:53-58.
(69) Kubo, T. and Y. Iwasa, 1996. Phenological pattern of tree regeneration in a model for forest species diversity. Theoretical Population Biology 49:90-117.
(68) Iwasa, Y., T. Kubo and K. Sato, 1995. Maintenance of forest species divesity and latitudinal gradient. Vegetatio 121:127-134.
(67) Pomiankowski, A. and Y. Iwasa, 1995. What causes diversity in male sexual characters ? Revue Swiss de Zoology 102:883-894.
(66) Kubo, T and Y. Iwasa, 1995. Inferring the rates of branching and extinction from molecular phylogenies. Evolution 49:694-704.
(65) Iwasa, Y. and A. Pomiankowski, 1995. Continual change in mate preferences. Nature 377:420-422.
(64) Harada, Y., H. Ezoe, Y. Iwasa, H. Matsuda and K. Sato, 1995. Population persistence and spatially limited social interaction. Theoretical Population Biology 48:65-91.
(63) Iwasa, Y., T.J. de Jong and P.G.L. Klinkhamer, 1995. Why pollinators visit only a fraction of the open flowers on a plant and its consequence for fitness curves of plants. Journal of Evolutionary Biology 8:439-453.
(62) Antonovics, J., Y. Iwasa and N.P. Hassell, 1995. A generalised model of parasitoid, veneral, and vector-based transmission processes. American Naturalist 145:661-675.
(61) Haccou, P. and Y. Iwasa, 1995. Optimal mixed strategies in stochastic environments. Theoretical Population Biology 47:212-243.
(60) Iwasa, Y. and S.A. Levin, 1995. The timing of life history events. Journal of Theoretical Biology 172:33-42.
(59) Yamauchi, A. and Y. Iwasa, 1995. Coupling of fermentation and foraging strategies of herbivorous mammals. Journal of Theoretical Biology 172:1-11.
(58) Iwasa, Y. and T. Kubo, 1995. Forest gap dynamics with partically synchronized disturbances and gap age distribution. Ecological Modelling 77:257-271.
(57) Harada, Y. and Y. Iwasa, 1994. Lattice population dynamics for plants with dispersing seeds and vegetative propagation. Researches on Population Ecology 36:237-249.
(56) Iwasa, Y. and A. Pomiankowski, 1994. Evolution of mate preferences for multiple sexual ornaments. Evolution 48:853-867.
(55) Iwasa, Y. and P. Haccou. 1994. ESS emergence pattern of male butterflies in stochastic environments. Evolutionary Ecology 8: 503-523.
(54) Ezoe, H., Y. Iwasa and T. Umeda, 1994. Aggregation by chemotactic random walk: drifting clusters and chemotactic friction. Journal of Theoretical Biology 168:259-267.
(53) Iwasa, Y. 1993. Overdispersed molecular evolution in constant environments. Journal of Theoretical Biology 164:373-393.
(52) Pomiankowski, A. and Y. Iwasa, 1993. Evolution of multiple sexual ornaments by Fisher's process of sexual selection. Proceeding of Royal Society, London, series B 253:173-181.
(51) Sato, K. and Y. Iwasa, 1993. Modelling of wave regeneration in subalpine Abies forests. Ecology 74:1538-1550.
(50) Kawano, K. and Y. Iwasa, 1993. A lattice-structured model for beech forest dynamics: the effect of understory dwarf bamboo. Ecological Modelling 66:261-275.
(49) Cohen, J.E., Y. Iwasa, Gh. Rautu, M. B. Ruskai, E. Seneta and Gh. Zbaganu. 1993. Relative entropy under mappings by stochastic matrices. Linear Algebra and Its Applications 179:211-235.
(48) Iwasa, Y., A. Yamauchi and S. Nozoe, 1992. Optimal seasonal timing of univotline and bivoltine insects. Ecological Research 7:55-62.
(47) Iwasa, Y. 1991. Asynchronous pupation of univoltine insects as the evolutionarily stable phenology. Researches on Population Ecology 33:213-227.
(46) Iwasa, Y. 1991. Pessimistic plants: the optimal growth schedule in fluctuating environments. Theoretical Population Biology 40:246-268.
(45) Iwasa, Y., K. Sato, and S. Nakashima, 1991. Dynamic modelling of wave regeneration (Shimagare) in subalpine Abies forests. Journal of Theoretical Biology 152:143-158.
(44) Iwasa, Y., A. Pomiankowski and S. Nee. 1991. The evolution of costly mate preferences. II. The "handicap" principle. Evolution 45:1431-1442.
(43) Pomiankowski, A., Y. Iwasa and S. Nee. 1991. The evolution of costly mate preferences. I. Fisher and biased mutation. Evolution 45:1422-1430.
(42) Iwasa, Y. 1991. Sex change evolution and cost of reproduction. Behavioral Ecology 2:56-58.
(41) Sasaki, A. and Y. Iwasa. 1991. Optimal growth schedule of pathogens within a host: switching between lytic and latant cycles. Theoretical Population Biology 39:201-239.
(40) Iwasa, Y. 1990. Evolution of selfing rate and resource allocation models. Plant Species Biology 5:19-30.
(39) Cuthill, I., A. Kacelnik, J.R. Krebs, P. Haccou, and Y. Iwasa, 1990. Starlings exploiting patches: the effect of recent experience on foraging decisions. Animal Behaviour 40:625-640.
(38) Iwasa, Y. and Y. Obara, 1989. A game model for the daily activity schedule of the male butterfly. Journal of Insect Behavior 2:589-608.
(37) Iwasa, Y., and D. Cohen, 1989. Optimal growth schedule of a perennial plant. American Naturalist 133:480-505.
(36) Iwasa, Y., S.A. Levin, and V. Andreasen, 1989. Aggregation in model ecosystems. II. Approximate aggregation. IMA Journal of Mathematics Applied in Medicine and Biology 6:1-23.
(35) Ishii, K., H. Matsuda, Y. Iwasa, and A. Sasaki, 1989. Evolutionarily stable mutation rate in a periodically fluctuating environment. Genetics 121:163-174.
(34) Iwasa, Y. 1988. Free fitness that always increases in evolution. Journal of Theoretical Biology 135:265-281.
(33) Iwasa, Y. and H. Mochizuki, 1988. Probability of population extinction accompanying a temporary decrease of population size. Researches on Population Ecology 30:145-164.
(32) Odendaal, F.J. and Y. Iwasa, 1987. Operational sex ratio in Ranidella signifera and Ranidella riparia (Anura: Leptodactylidae). Copeia 1987:1025-1928.
(31) Iwasa, Y., V. Andreasen, and S.A. Levin, 1987. Aggregation in model ecosystems. I. Perfect aggregation. Ecological Modelling 37:287-302.
(30) Sasaki, A. and Y. Iwasa, 1987. The optimal recombination rate in fluctuating environments. Genetics 115:377-388.
(29) Iwasa, Y. and A. Sasaki, 1987. Evolution of the number of sexes. Evolution 41:49-65.
(28) Takada, T. and Y. Iwasa, 1986. Size distribution dynamics of plants with interation by shading. Ecological Modelling 33:173-184.
(27) Iwasa, Y. and J. Roughgarden, 1986. Interspecific competition among metapopulations with space-limited subpopulations. Theoretical Population Biology 30:194-214.
(26) Levin, S.A., M. A. Harwell, and 11 contributors, 1986. Potential ecological consequences of genetically engineered organisms. Environmental Management 10:495-513.
(25) Roughgarden, R. and Y. Iwasa, 1986. Dynamics of a metapopulation with space-limited subpopulations. Theoretical Population Biology 29:235-261.
(24) Liu, W-M., S.A. Levin, and Y. Iwasa, 1986. Influence of nonlinear incidence transmission rates upon the behavior of SIRS epidemic models. Journal of Mathematical Biology 23:187-204.
(23) Iwasa, Y. and J. Roughgarden, 1985. Evolution in a metapopulation with space-limited subpopulations. IMA Journal of Mathematics Applied in Medicine and Biology 2:93-107.
(22) Odendaal, F.J., Y. Iwasa, and P.R. Ehrlich, 1985. Duration of female availability and its effect on butterfly mating systems. American Naturalist 125:673-678.
(21) Roughgarden, J., Y. Iwasa, and C. Baxter. 1985. Demographic theory for an open population with space-limited recruitment. Ecology 66:54-67.
(20) Iwasa, Y., D. Cohen, and J.A. Leon. 1985. Tree height and crown shape as result of competitive games. Journal of Theoretical Biology 112:279-297.
(19) Iwasa, Y., Y. Suzuki, and H. Matsuda. 1984. Theory of oviposition strategy of parasitoids. I. Effect of mortality and limited egg number. Theoretical Population Biology 26:205-227.
(18) Kakehashi, M., Y. Suzuki, and Y. Iwasa. 1984. Niche overlap of parasitoids in host-parasitoid systems: its consequence to single vs. multiple introduction controversy in biological control. Journal of Applied Ecology 21:115-131.
(17) Iwasa, Y. and F.J. Odendaal. 1984. A theory on the temporal pattern of operational sex ratio during the breeding season: the active-inactive model. Ecology 65:886-893.
(16) Iwasa, Y. and E. Teramoto. 1984. Branching-diffusion model of the formation of a population's distributional pattern. Journal of Mathematical Biology 19:109-124.
(15) Iwasa Y. and J. Roughgarden. 1984. Shoot/root balance of plants: Optimal growth of a system with many vegetative organs. Theoretical Population Biology 25:78-105.
(14) Iwasa, Y., F. Odendaal, D. Murphy, P. Ehrlich, and A. Launer. 1983. Emergence patterns in male butterflies: A hypothesis and a test. Theoretical Population Biology 23:363-379.
(13) Iwasa, Y. 1982. Vertical migration of zooplankton: A game between predator and prey. American Naturalist 120:171-180.
(12) Ito, Y. and Y. Iwasa. 1981. Evolution of litter size. I. Conceptual reexamination. Researches on Population Ecology 23:344-359.
(11) Iwasa, Y. 1981. Role of sex ratio in the evolution of eusociality in haplodiploid social insects. Journal of Theoretical Biology 93:125-142.
(10) Higashi, M., T. Miura, K. Tanimizu, and Y. Iwasa. 1981. Effect of the feeding activity of snails on the biomass and productivity of an algal community attached to a reed stem. Verh. Internat. Verein. Limnol. 21:558-563.
(9) Iwasa, Y. 1981. Size distribution in a biological population: A theoretical analysis. Verh. Internat. Verein. Limnol. 21:1304-1308.
(8) Iwasa, Y., M. Higashi, and N. Yamamura. 1981. Prey distribution as a factor determining the choice of optimal foraging strategy. American Naturalist 117:710-723.
(7) Suzuki, Y. and Y. Iwasa. 1980. A sex ratio theory of gregarious parasitoids. Researches on Population Ecology 22:366-382.
(6) Iwasa, Y. and E. Teramoto. 1980. A criterion of life history evolution based on density dependent selection. Journal of Theoretical Biology 84:545-566.
(5) Iwasa, Y. 1978. Optimal death strategy of animal populations. Journal of Theoretical Biology 72:611-626.
(4) Miura, T., K. Tanimizu, Y. Iwasa, and A. Kawakita. 1978. Macroinvertebrates as an important supplier of nutrients in a dense macrophyte zone in Lake Biwa. Vehr. Internat. Verin. Limnol. 20:1116-1121.
(3) Iwasa, Y. and E. Teramoto. 1977. A mathematical model for the formation of a distributional pattern and an index of aggregation. Japanese Journal of Ecology 27:117-124.
(2) Miura, T., A. Kawakita, Y. Iwasa, and K. Tanimizu. 1976. Studies on the submerged plant community in Lake Biwa. II. Macroinvertebrates as an important supplier of nitrogeneous nutrients in a dense macrophyte zone. Physiology and Ecology, Japan 17:589-593.
(1) Iwasa, Y. and T. Miura. 1976. The rate of respiration and ammonia-excretion of freshwater shrimps in Lake Biwa. Physiology and Ecology, Japan 17:465-472.

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